Ok - so I had a call with the Guru and he offered some more suggestions for things to check: namely:
1: try rotating the engine with just one push-rod in, see if it's one or the other causing the issue.
2: check the play of the camshaft in it's bushes with no springs (ie whip the pushrods out)
3: measure the wear on the bearing parts of the camshaft spindle (ie where it sits in the bushes.
So - I did those.
1: Each "side" of the timing still has the click noises, but with only one valve in operation I was able to figure out exactly where in the rotation the clicks are happening. BOTH of them are about 5 degrees after peak cam lift. IE when the cam-follower has "just" passed the highest point of opening on the cam lobe.
2: there is f***-all play in the camshaft when the timing cover is in place, at all 4 cardinal points (TDC, TDC+90, BDC, BDC+90) whatever play there is, is identical. With a nadger of end-play. I think the bushes are fine.
3: I measured the cam spindle-ends with a metric vernier, then converted to imperial (see last photo in the post), to get the following:
crank end: 0.003071 under (nominal is 0.5", measurements avg to 0.496929")
mag end: 0.001969 under (nominal 0.5", measurements avg to 0.498031")

I also measured the mag-end bush inner diameter (the one in the timing cover): 0.497"

(I haven't got any means of measuring the crank-end bush without stripping the engine again, and even then I'd only be measuring the outer end of the bush, relative to the spindle).

So, If the bushes are fine, and even though there's a touch of end-play on the cam-followers, there's no other movement on that spindle (you need SOME play to prevent the followers binding on each other), ie both followers are not rattling on the spindle: we are both stumped.
Even the Guru is clutching at straws now: his last communique was an idea to check the actual cam-lobe for a slight dip at the point the click is happening. IE use a dial guage to measure each lobe as it rotates (preferably in a lathe, but I should be able to bodge up something reasonably solid....)
The really weird thing about this is that the click noise happens even when the crank is being rotated
really slowly, so the thought that it's a "bounce" from the momentum of the parts in the system doesn't really add up.
My current thought is to say F**k it and just finish the build and see what happens! getting bored now...
(as an aside, replacing the camshaft bushes sounds like a right faff... the one in the timing cover needs to be machined after fitting to give the correct end play, and they need to be reamed to size "inline", ie both fitted in place when the engine is completely torn down. I'm quite glad I don't need to do that!)